Ecology and Evolutionary Biology
A308 Earth and Marine Sciences
(831) 459-5358
http://www.eeb.ucsc.edu
Physical and Biological Sciences Undergraduate Affairs Office
142 Jack Basking Engineering Bldg.
(831) 459-4143
http://undergrad.pbsci.ucsc.edu
EEB Faculty | EEB Program Statement
Lower-Division Courses
20B. Development and Physiology. F,W,S
Topics in morphology, physiology, development, genetics, and endocrinology selected to exemplify current issues and perspectives in organismic biology. Prerequisite(s): BIOL 20A. (F) M. Peterson, (WS) R. Dunkin
20C. Ecology and Evolution. F,W,S
Introduction to ecology and evolution covering principles of evolution at the molecular, organismal, and population levels. Evolutionary topics include genetic and phenotypic variation, natural selection, adaptation, speciation, and macroevolution. Also covers behavioral, population, and community ecology including applied ecological issues. B. Marinovic
75. Scientific Diving Certification (2 credits). F,S
Prerequisite for course 161/L, Kelp Forest Ecology, and all research diving performed under the auspices of UCSC or other academic institutions. Course work includes lectures and scuba diving. Topics include subtidal sampling techniques, navigation, low visibility diving, search and recovery, rescues, small boat use, oxygen administration for divers, technical blue water deep diving, physics, and physiology. Apply online at http://www2.ucsc.edu/sci-diving. Students are billed a course materials fee that covers costs for equipment use, materials, and transportation. Prerequisite(s): skill level equal to Advanced Scuba Diver Certification, pass scuba physical, provide own scuba gear, be certified in CPR and First Aid; and interview: pass swim test and scuba skills test. Enrollment limited to 16. S. Clabuesch
80S. Lies, Damn Lies, and Statistics. *
Probability and statistics underlie much of our everyday experience and, as such, there is a fundamental need for an understanding of the use, and misuse, of statistics. This course is taught through case studies based in biology, politics, economics, crime, education, disease, conservation, and other fields of interest. For example, does a change in crime rate really affect your probability of being a victim of a crime? The goal is to provide all students with sufficient understanding probability and statistics to determine if everyday and often sensationalistic reporting of "statistical" results is meaningful. (General Education Code(s): SR.) P. Raimondi
85. Natural History of the UCSC Natural Reserves. *
Lectures and field trips familiarize students with the flora and fauna of the UCSC Natural Reserves. Field trips focus on surveying and identifying vertebrates and plants at each UCSC Natural Reserve (Fort Ord, Campus Reserve, Big Creek, Younger Lagoon, and Ano Nuevo). (General Education Code(s): PE-E.) The Staff
95. Seymour Center Docent Training (2 credits). W
Taught as a series of seminars, course provides a survey of marine sciences and the role of scientific research in understanding and conserving the world's oceans. Topics include: marine biology, ecology, conservation, coastal geology, and climate change. This series is intended to prepare students to interpret research and inform the public by leading tours at the Seymour Marine Discovery Center at the Long Marine Lab. Enrollment by application and interview. (General Education Code(s): PR-S.) M. Carr
99. Tutorial. F,W,S
Individual, directed study for undergraduates. Students submit petition to sponsoring agency. May be repeated for credit. The Staff
Upper-Division Courses
107. Ecology. W,S
Focuses on physiological, behavioral, and population ecology, and on linking ecological processes to evolution. It includes basic principles, experimental approaches, concepts of modeling, and applications to ecological problems. Prerequisite(s):satisfaction of the Entry Level Writing and Composition requirements; BIOL 20A, BIOE 20B, and BIOE 20C. (W) B. Lyon, (S) J. Estes
108. Marine Ecology. W
Paradigms and designs in marine ecology. A review of the paradigms that have shaped our understanding of marine ecology; analysis and discussion of experiments with these paradigms. Students cannot receive credit for this course and course 208. Prerequisite(s): satisfaction of the Entry Level Writing and Composition requirements; BIOL 20A, BIOE 20B, and BIOE 20C; BIOE 107 or 140 recommended. Enrollment restricted to juniors and seniors. M. Carr, P. Raimondi
109. Evolution. F,W
An examination of the history and mechanisms of evolutionary change. Topics include molecular evolution, natural and sexual selection, adaptation, speciation, biogeography, and macroevolution. Prerequisite(s): satisfaction of the Entry Level Writing and Composition requirements; BIOL 20A, BIOE 20B, BIOE 20C, and BIOL 105. (F) K. Kay, (W) G. Pogson
112. Ornithology. *
Introduction to the evolution, ecology, behavior, and natural history of birds, using exemplary case histories to illustrate key concepts in evolution, ecology, and behavior. Prerequisite(s): BIOE 107, BIOE 109, or BIOE 140. Concurrent enrollment in BIOE 112L is required. B. Lyon
112L. Ornithology Field Studies (2 credits). *
Field trips introduce students to field identification skills and field investigation of census, foraging behavior, migration, social behavior, and communication. Examination of specimens in the laboratory will be used to highlight the diversity and taxonomy of birds. Students are billed a materials fee. Some field trips may require students to provide their own transportation. Prerequisite(s): BIOE 107, BIOE 109, or BIOE 140. Concurrent enrollment in BIOE 112 is required. Offered in alternate academic years. B. Lyon
114. Herpetology. *
Lectures introduce students to evolution, development, physiology, behavior, ecology, and life history of reptiles and amphibians. The materials integrate with conceptual and theoretical issues of ecology, evolution, physiology, and behavior. Prerequisite(s): BIOE 107, BIOE 109, BIOE 110, or BIOE 140. Concurrent enrollment in BIOE 114L required. Offered in alternate academic years. B. Sinervo
114L. Field Methods in Herpetological Research (2 credits). *
Field trips introduce students to natural history, censusing techniques, physiological ecology, and behavioral analysis of reptiles and amphibians. Laboratories introduce students to techniques for analyzing behavior and physiology. Field studies culminate with a group project in a natural setting. Some field trips may be held on weekends due to weather considerations. Some field trips may require students to provide their own transportation, some transportation will be provided by UCSC. Students are billed a materials fee. Prerequisite(s): BIOE 107, 109, 110, or 140. Concurrent enrollment in BIOE 114 is required. Offered in alternate academic years. B. Sinervo
117. Systematic Botany of Flowering Plants. W
An examination of the taxonomy and evolution of flowering plants. Special topics include phylogenetics and cladistics, plant species concepts, and modern methods of systematic research. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. Must be taken concurrently with BIOE 117L. K. Kay
117L. Systematic Botany of Flowering Plants Laboratory (2 credits). W
Weekly laboratory concerned primarily with California flora and plant families. Several field trips. Students are billed a materials fee. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. Must be taken concurrently with BIOE 117. K. Kay
118. Plants and Society: the Biology of Food, Shelter, and Medicine. S
Introduces plant biology as it affects human society. Topics include the origins of agriculture, the morphology and chemistry of food plants, the material uses of plant products, the biology of medicinal plants, and plant diversity and bioprospecting. Prerequisite(s): BIOL 20A and BIOE 20B; and BIOE 20C or ENVS 23 and ENVS 24 J. Pittermann, I. Parker
120. Marine Botany. S
An introduction to the biology of marine algae, fungi, and angiosperms with regard to form and function. Major boreal, temperate, and tropical marine plant communities. Lecture format. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. Must be taken concurrently with BIOE 120L. The Staff
120L. Marine Botany Laboratory (2 credits). S
One laboratory weekly and several field trips. Focuses on marine algae, fungi, and angiosperms. Students are billed a materials fee. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. Must be taken concurrently with BIOE 120. The Staff
122. Invertebrate Zoology. W
An examination of invertebrates and their habitats. Lecture format. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. BIOE 122L must be taken concurrently. B. Marinovic
122L. Invertebrate Zoology Laboratory (2 credits). W
An examination of invertebrates and their habitats. Weekly laboratories or field trips. Students are billed a materials fee. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. BIOE 122 must be taken concurrently. B. Marinovic
124. Mammalogy. F
Introduces the biology of mammals, including their classification, evolution, behavior, reproductive strategies, and general ecology. Examines the diagnostic traits of mammals; provides a survey of the living orders along with their diagnostic features, physiological and behavioral specializations, and adaptations. Prerequisite(s): BIOL 20A; and BIOE 20B and 20C. Concurrent enrollment in course 124L is required. The Staff
124L. Mammalogy Laboratory (2 credits). F
Focuses on the identification of mammals and their specific traits. Exercises provide hands-on experience at identifying mammal orders, families, and species. Field trip provides students with field techniques in mammalogy. Prerequisite(s): BIOL 20A; and BIOE 20B and 20C. Concurrent enrollment in course 124 is required. The Staff
127. Ichthyology. *
An introduction to the biology of jawless, cartilaginous, and bony fishes—their classification, evolution, form, physiology, and ecology. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. BIOE 127L must be taken concurrently. Offered in alternate academic years. G. Bernardi
127L. Ichthyology Laboratory (2 credits). *
One laboratory session a week and several field trips to study the biology of fish. Students are billed a materials fee. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. BIOE 127 must be taken concurrently. Offered in alternate academic years. G. Bernardi
128. Ecology and Physiology of Large Marine Vertebrates. S
Lectures and laboratory computer exercises familiarize students with research methods, study design, statistics, and research tools for large marine vertebrates (seals, birds, fish, and sharks). Research topics include: animal tracking; diving physiology; behavior; foraging ecology; and energetics. Prerequisite(s): BIOL 20A; and BIOE 20B and 20C. P. Robinson
128L. Large Marine Vertebrates Field Course. S
Lectures combined on fieldwork with large marine vertebrates in the laboratory and lectures with large marine vertebrates in the field (Monterey Bay, Ano Nuevo). Fieldwork familiarizes students with research methods, study design, and statistical approaches for research on large marine vertebrates (seals, birds, fish, and sharks). Research includes: animal tracking; physiology; behavior; foraging ecology; and energetics. Students are billed a materials fee. Prerequisite(s): BIOL 20A; and BIOE 20B, 20C, and 128. Enrollment limited to 24. P. Robinson
129. Biology of Marine Mammals. S
A survey of cetaceans, pinnipeds, sirenians, and sea otters, including natural history, systematics, physiology, behavior, anatomy, and conservation. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C; BIOL 110 is recommended. D. Costa
129L. Biology of Marine Mammals Laboratory (2 credits). S
Covers the basics of marine mammal taxonomy, anatomy, and field methods with an emphasis on local field identification and understanding of local species. Will include field trips to Long Marine Lab, Ano Nuevo, and Monterey Bay. Students are billed a materials fee. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. Must be taken concurrently with BIOE 129. D. Costa
131. Animal Physiology. W
Principles and concepts underlying the function of tissues and organ systems in animals with emphasis on vertebrate systems. Students cannot receive credit for this course and BIOL 130. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. R. Mehta, T. Williams
131L. Animal Physiology Laboratory (2 credits). W
Experiments conducted with primary focus on quantitative physiological principles of organ systems and intact organisms. Students cannot receive credit for this course and course 130L. Students are billed a materials fee. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. Concurrent enrollment in BIOE 131 is required. R. Mehta, T. Williams
133. Exercise Physiology. S
An advanced-level course concerning physiological and biochemical processes associated with human performance. Emphasis is on the integration of organ systems for exercise. Topics include metabolism and fuel utilization, cardiovascular and respiratory dynamics during activity, and the effects of training. Requires a good understanding of basic physiological function and anatomy. Prerequisite(s): BIOL 20A, BIOE 20B and 20C. BIOE 131 recommended. Concurrent enrollment in BIOE 133L required. Offered in alternate academic years. T. Williams
133L. Exercise Physiology Laboratory (2 credits). S
An introduction to basic measurement techniques used in assessing the physiological response of humans to exercise. Sessions cover oxygen consumption, respiratory rate, and heart rate monitoring during aerobic and anaerobic activity. Students are billed a materials fee. Prerequisite(s): BIOL 20A, and BIOE 20B and 20C. BIOE 131 recommended. Concurrent enrollment in BIOE 133 is required. Offered in alternate academic years. T. Williams
134. Comparative Vertebrate Anatomy. F
Course focuses on vertebrate form and function: an integration of physiology and biomechanics. Topics include: the physiology and biomechanics underlying vertebrate locomotion; vertebrate feeding; and the morphological changes associated with different locomotion and feeding strategies through evolutionary time. Prerequisite(s): BIOL 20A, BIOE 20B and BIOE 20C; Physics 6A. Concurrent enrollment in BIOE 134L is required. R. Mehta
134L. Comparative Vertebrate Anatomy Laboratory (2 credits). F
Course focuses on the gross dissections all major clades of vertebrates: development, form, and diversity of organ systems and basic principles of evolution; vertebrate classification; and functional morphology, with emphasis on feeding and locomotion. Anatomical dissections integrated with the associated lecture material focusing on biomechanics, form, and function. Students are billed for a materials fee. Prerequisite(s): BIOL 20A, BIOE 20B and BIOE 20C; Physics 6A. Concurrent enrollment in BIOE 134 is required. R. Mehta
135. Plant Physiology. W
Cellular and organismal functions important in the life of green plants. Prerequisite(s): BIOL 20A and BIOE 20B and 20C; concurrent enrollment in course 135L is required. J. Pittermann
135L. Plant Physiology Laboratory (2 credits). W
Weekly laboratory concerning the cellular and organismal functions of green plants. Students are billed a materials fee. Prerequisite(s): BIOL 20A and BIOE 20B and BIOE 20C; concurrent enrollment in course 135. J. Pittermann
137. Molecular Ecology. W
This combination lecture/laboratory course explores the use of molecular (DNA and/or protein) data in ecological and conservation research. Topics covered include data collection; marker choice; estimating genetic diversity and population structure; the inference of mating systems; and environmental genomics. Prerequisite(s): courses 20B and 20C and BIOL 20A and BIOL105 Concurrent enrollment in course 137L is required. Enrollment limited to 24. B. Shapiro
137L. Molecular Ecology Laboratory (2 credits). W
This combination lecture/laboratory course explores the use of molecular (DNA and/or protein) data in ecological and conservation research. Topics covered include data collection; marker choice; estimating genetic diversity and population structure; the inference of mating systems; and environmental genomics. Prerequisite(s): courses 20B and 20C and BIOL 20A and BIOL105. Concurrent enrollment in course 137 is required. Enrollment limited to 24. B. Shapiro
140. Behavioral Ecology. F
An introduction to social and reproductive behavior. Emphasis on studies of vertebrates in their natural habitat. Ideas concerning the evolution of social behavior, mating systems, and individual reproductive strategies. Case histories of well-studied animals that illustrate key principles in courtship and mating, parental behavior, and food-getting behavior. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. B. Sinervo
141L. Behavioral Ecology Field Course. W
A field-based course introducing students to concepts and methods for studying behavioral ecology in nature. Students will conduct observations and field experiments on various local model organisms including elephant seals, hummingbirds, sparrows, lizards, ants, bees, frogs, and salamanders. Students are billed a materials fee. Prerequisite(s): BIOE 107 or BIOE 140 or BIOE 110; satisfaction of the Entry Level Writing and Composition requirements. Enrollment limited to 25. Offered in alternate academic years. (General Education Code(s): W.) B. Sinervo, B. Lyon
145. Plant Ecology. F
An exploration of the ecology of plant form, function, distribution, abundance, and diversity. Topics include plant adaptations to environmental conditions, life history variation, competition, reproductive ecology, herbivory, and patterns of diversity. Lecture with discussions of original papers and independent field project. Students cannot receive credit for this course and course 245. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. BIOE 107 is recommended. I. Parker
145L. Field Methods in Plant Ecology. F
Hands-on exploration of the concepts and techniques of plant ecology. A combination of lab, greenhouse, and field-based exercises (irrespective of weather conditions). Statistical analysis and scientific writing. One required weekend field trip. Students cannot receive credit for this course and course 245L. Students are billed a materials fee. Prerequisite(s): satisfaction of the Entry Level Writing and Composition requirements; BIOL 20A, BIOE 20B, and BIOE 20C. Concurrent enrollment in BIOE 145 is required. BIOE 107 is recommended. (General Education Code(s): W.) I. Parker
147. Community Ecology. S
Develops the major themes of community biology: structure, trophic dynamics, succession, complex interactions among species, herbivory, evolution and coevolution. Uses case histories of well-studied marine and terrestrial systems. Students cannot receive credit for this course and course 247. Prerequisite(s): BIOE 107, 108, 145, 155 or 159A; or Environmental Studies 24 by permission of instructor. L. Fox
148. Quantitative Ecology. *
Quantitative treatment of the central concepts and applications of theoretical ecology. Emphasis on the mathematical modeling of single populations and species interactions, and the integration of models with data. Topics include stochastic and deterministic processes of extinction; discrete- and continuous-time models of growth and population viability analysis relevant to small and harvested populations; numeric and analytical investigations of dynamics and stability; introduction to model-fitting in information theoretic framework using R and/or MATLAB. Prerequisite(s): BIOE 107. M. Tinker
149. Disease Ecology. S
Focuses on the ecological and evolutionary processes that drive the transmission of pathogens between hosts; the impact of disease on host populations; and what causes the emergence of an infectious disease. Includes theoretical framework, description of field techniques, and discussion of wildlife and human diseases including malaria, West Nile virus, Lyme disease, HIV, avian influenza (bird flu), Chikungunya, tuberculosis, chytridiomycosis, and Ebola. Prerequisite(s): BIOL 20A, and BIOE 20B and 20C and 107. A. Kilpatrick
150. Ecological Field Methods. S
Lectures and laboratory computer exercises designed to familiarize students with research methods, study design, statistical approaches, and analysis tools for ecological research. Students cannot receive credit for this course and Environmental Studies 104A. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C; concurrent enrollment in BIOE 150L is required. BIOE 107, 108, 140, or 147 recommended. Enrollment limited to 25. D. Croll
150L. Ecological Field Methods Laboratory. S
Field-oriented course in the study of animal ecology and behavior. Combines overview of methodologies and approaches to field research with practical field studies. Students are billed a materials fee. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C; concurrent enrollment in BIOE 150 is required. BIOE 107, 108, 140, or 147 recommended. Enrollment limited to 25. (General Education Code(s): W.) D. Croll
151A. Ecology and Conservation in Practice Supercourse: Ecological Field Methods. *
An intensive, on-site learning experience in terrestrial field ecology and conservation, using the University of California Natural Reserves. Students study advance concepts in ecology, conservation, and field methods for four weeks, then experience total immersion in field research at the UC Natural Reserves. Lectures, field experiments, and computer exercises familiarize students with research methods, study design, statistical approaches, and analytical tools for ecological research. Enrollment by application. Prerequisite(s): BIOL 20A, BIOE 20B, BIOE 20C or ENVS 23, 24, 100; and AMS 7 and 7L. Concurrent enrollment in BIOE 151B-C-D or ENVS 109B-C-D is required. Satisfies the senior exit requirement for biological sciences majors and satisfies the senior exit requirement for environmental studies majors by prior approval. Students cannot receive credit for this course and BIOE 150, 150L, ENVS 104A or 196A. (Also offered as Environmental Studies 109A. Students cannot receive credit for both courses.) D. Croll, E. Zavaleta
151B. Ecology and Conservation in Practice Supercourse: Ecological Field Methods Laboratory. *
Field-oriented course in ecological research. Combines overview of methodologies and approaches to field research with practical field studies. Students complete field projects in ecology and also learn the natural history of the flora and fauna of California. Students are billed a materials fee. Enrollment by application. Prerequisite(s): Entry Level Writing and Composition requirements; BIOL 20A, BIOE 20B, BIOE 20C or ENVS 23, 24, 100; and AMS 7 and 7L. Concurrent enrollment in BIOE 151A-C-D or ENVS 109A-C-D is required. Satisfies the senior exit requirement for biological sciences majors and satisfies the senior exit requirement for environmental studies majors by prior approval. Students cannot receive credit for this course and BIOE 150, 150L, ENVS 104A or 196A. (Also offered as Environmental Studies 109B. Students cannot receive credit for both courses.) (General Education Code(s): W.) D. Croll, E. Zavaleta
151C. Ecology and Conservation in Practice Supercourse: Functions and Processes of Terrestrial Ecosystems. *
From lectures and discussion of terrestrial community and ecosystem ecology, students work individually or in small groups to present an idea for a project, review relevant literature, develop a research question/hypothesis, design and perform an experiment, collect and analyze data, and write a report. The instructor evaluates the feasibility of each student's project before it begins. Enrollment by application. Prerequisite(s): BIOL 20A, BIOE 20B, BIOE 20C or ENVS 23, 24, 100; and AMS 7 and 7L. Concurrent enrollment in BIOE 151A-B-D or ENVS 109A-B-D is required. Satisfies the senior exit requirement for biological sciences majors and satisfies the senior exit requirement for environmental studies majors by prior approval. Students cannot receive credit for this course and BIOE 150, 150L, ENVS 104A or 196A. (Also offered as Environmental Studies 109C. Students cannot receive credit for both courses.) D. Croll, E. Zavaleta
151D. Ecology and Conservation in Practice Supercourse: Conservation in Practice. *
Focuses on current issues in environmental and conservation biology and the emerging field methods used to address them. From field-oriented lectures about current issues in environmental and conservation biology, students pursue research project as individuals and small groups to develop hands-on experience with field skills in conservation research and resource management. Enrollment by application. Prerequisite(s): BIOL 20A, BIOE 20B, BIOE 20C or ENVS 23, 24, 100; and AMS 7 and 7L. Concurrent enrollment in BIOE 151A-B-C or ENVS 109A-B-C is required. Satisfies the senior exit requirement for biological sciences majors and satisfies the senior exit requirement for environmental studies majors by prior approval. Students cannot receive credit for this course and BIOE 150, 150L, ENVS 104A or 196A. (Also offered as Environmental Studies 109D. Students cannot receive credit for both courses.) D. Croll, E. Zavaleta
155. Freshwater Ecology. F
Provides an overview of the physical, chemical, and biological processes that characterize inland waters such as lakes, streams, rivers, and wetlands. Also addresses relationships between humans and freshwater, and discusses these challenges in conservation. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. E. Palkovacs
155L. Freshwater Ecology Laboratory. S
Field and laboratory study of the ecology of freshwater systems including lakes, streams, and estuaries. Students gain experience sampling and identifying freshwater organisms, designing and analyzing ecological experiments, and writing scientific reports. Enrollment limited to 24. E. Palkovacs
158L. Marine Ecology Laboratory. *
Supervised individual research projects in experimental marine biology. Students carry out a complete research project, including (1) the formation of hypotheses; (2) the design and implementation of experiments; (3) collection, analysis, and interpretation of data; and (4) write-up of an oral presentation. Students are billed a materials fee. Prerequisite(s): BIOE 108; satisfaction of the Entry Level Writing and Composition requirements. Offered in alternate academic years. (General Education Code(s): W.) M. Carr, P. Raimondi
159A. Marine Ecology Field Quarter: Marine Ecology with Laboratory. F
Total immersion in marine ecology for very motivated students. Students develop a research project during first five weeks on campus and then spend five weeks of immersion in directed research without distraction in isolated locations off campus (past locations include the Gulf of California in Mexico and Moorea in French Polynesia). Not available through University Extension. No other courses may be taken during this quarter. Students must sign a contract agreeing to standards of behavior outlined in the UCSC Rule Book and by the instructors. Students are billed a materials, transportation (not airfare), and room and board fee. Paradigms and designs in marine ecology. A review of the paradigms that have shaped our understanding of marine ecology and analysis and discussion of experiments with these paradigms. Students carry out a complete research project, including the formation of hypotheses; the design and implementation of experiments; the collection, analysis, and interpretation of data; and the write-up and oral presentation of results. Admission by interview during previous winter quarter. BIOE 159A, 159B, 159C, and 159D are equivalent to BIOE 127, 127L, 108, and 158L for major requirements. Prerequisite(s): satisfaction of the Entry Level Writing and Composition requirements; BIOE 159A, 159B, 159C, and 159D must be taken concurrently. Enrollment limited to 26. Offered in alternate academic years. (General Education Code(s): W.) P. Raimondi, G. Bernardi
159B. Marine Ecology Field Quarter: Ichthyology with Laboratory. F
An introduction to the biology of jawless, cartilaginous, and bony fishes—their classification, evolution, form, physiology, and ecology. Admission by interview during previous winter quarter. BIOE 159A, 159B, 159C, and 159D are equivalent to BIOE 127, 127L, 108, and 158L for major requirements. BIOE 159A, 159B, 159C, and 159D must be taken concurrently. Enrollment limited to 26. Offered in alternate academic years. P. Raimondi, G. Bernardi
159C. Marine Ecology Field Quarter: Methods in Field Ecology. F
Students learn quantitative methods for field experiments and surveys. Emphasis will be on marine environments, but there will also be exposure to terrestrial systems. This is the lecture component to course 159D. No text is required for this course; instead, readings from the current literature will be assigned. Students are evaluated on written independent field project proposals and class participation. Admission by interview during previous winter quarter. BIOE 159A, 159B, 159C, and 159D are equivalent to BIOE 127, 127L, 108, and 158L for major requirements. BIOE 159A, 159B, 159C, and 159D must be taken concurrently. Enrollment limited to 26. Offered in alternate academic years. P. Raimondi, G. Bernardi
159D. Marine Ecology Field Quarter: Methods in Field Ecology Laboratory. F
This is laboratory portion of course 159C. Students carry out independent field projects under the supervision of course instructors. All work is done during the 5–6 week off-campus portion of course 159. Students are evaluated on field techniques, the final write-up of their independent field projects, and class participation. Admission by interview during previous winter quarter. BIOE 159A, 159B, 159C, and 159D are equivalent to BIOE 127, 127L, 108, and 158L for major requirements. BIOE 159A, 159B, 159C, and 159D must be taken concurrently. Enrollment limited to 26. Offered in alternate academic years. P. Raimondi, G. Bernardi
161. Kelp Forest Ecology. *
Study of organization of kelp forests as models for examining biological communities. The physical and biotic factors responsible for community organization of kelp forests are explored using original literature and data collected in BIOE 161L. Class meets one full morning each week. Prerequisite(s): by interview only; BIOL 20A, BIOE 20B, and BIOE 20C are required. Students must pass the University Research Diving Certification (contact the diving safety officer, Institute of Marine Sciences, for further information). Enrollment restricted to seniors. BIOE 161L must be taken concurrently; BIOE 107, 120/L, 122/L are recommended. Enrollment limited to 24. Offered in alternate academic years. M. Carr, P. Raimondi
161L. Kelp Forest Ecology Laboratory. *
Fieldwork using SCUBA to quantitatively and qualitatively examine the abundance and distribution of organisms in kelp forests, with additional laboratory work. Culminates with a directed individual research project. Class meets one full morning each week. Students are billed a materials fee. Admission by interview. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C; satisfaction of the Entry Level Writing and Composition requirements; BIOE 161 must be taken concurrently; BIOE 107, 120/L, 122/L are recommended. Students must pass the University Research Diving Certification (contact the Diving Safety Officer, Institute of Marine Sciences, for further information). Enrollment limited to 24. Offered in alternate academic years. (General Education Code(s): W.) M. Carr, P. Raimondi
163. Ecology of Reefs, Mangroves, and Seagrasses. W
Integrated treatment of coral reefs, sea grasses, and mangroves emphasizing interactions and processes through time. Major topics: biological and geological history, biogeography, evolution and ecology of dominant organisms, biodiversity, community and ecosystem ecology, geology, biogeochemistry, global change, human impacts. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. Concurrent enrollment in BIOE 163L is required. D. Potts
163L. Ecology of Reefs, Mangroves, and Seagrasses Laboratory (2 credits). W
An interdisciplinary laboratory exploration of the anatomy, morphology, adaptations, diversity, evolution, and ecology of corals, mangroves, and seagrasses and of their physical, chemical, and geological environments. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C. Concurrent enrollment in BIOE 163 is required. D. Potts
165. Marine Conservation Biology. F
Initially undertakes an in-depth comparison of the biology and conservation of marine versus terrestrial ecosystems. With this foundation, course examines marine biodiversity loss resulting from overexploitation, habitat loss, species introduction, and pollution, with particular emphasis on the resulting trophic cascades, biodiversity losses, and climate change. Students cannot receive credit for this course and Environmental Studies 120. Prerequisite(s): BIOL 20A, BIOE 20B, and BIOE 20C; OCEA 101 recommended. D. Croll
172. Population Genetics. F
Basic population genetics and selected topics will be covered, including genetics of speciation, tempo and mode of evolution, genetics of social behavior, natural selection in human populations, and the impact of molecular studies on evolutionary theory. Students cannot receive credit for this course and BIOE 272. Prerequisite(s): BIOL 20A, BIOE 20B, BIOE 20C, and BIOL 105. Concurrent enrollment in BIOE 172L is required. Offered in alternate academic years. G. Pogson
172L. Population Genetics Laboratory (2 credits). F
A companion course to 172, Population Genetics, that applies the theory developed in that course to related disciplines including conservation biology, ecology, agriculture, and population biology. Original scientific literature relating to the theory developed in BIOE 172 is read, and applied problem sets are solved by the students. Students cannot receive credit for this course and BIOE 272L. Prerequisite(s): BIOL 20A, BIOE 20B, BIOE 20C, and BIOL 105. Concurrent enrollment in BIOE 172 is required. Offered in alternate academic years. G. Pogson
182F. Exploring Research in EEB (2 credits). *
Provides undergraduate students with exposure to research in the laboratory of an Ecology and Evolutionary Biology (EEB) faculty member, affiliate, or adjunct. Students are not expected to do independent research but rather to assist in laboratory or field research projects under the supervision of the faculty mentor or appointed researcher. Prerequisite(s): Undergraduate research contract on file with the department. M. Carr
183L. Undergraduate Research in EEB (3 credits). *
Designed to ensure that students are intellectually engaged in the planning or implementation of a supervised or independent research project, achieve a fundamental understanding of implementing the scientific method, and develop their scientific writing and and presentation skills. Prerequisite(s): concurrent enrollment in course 183W and an Undergraduate Research Contract on file with the department. (General Education Code(s): W satisfied by taking this course and course 183W.) The Staff
183W. Undergraduate Research in EEB--Writing (2 credits). F,W,S
Ensures that students are intellectually engaged in the planning or implementation of a supervised or independent research project, achieve a fundamental understanding of implementing the scientific method, and develop their scientific writing and presentation skills. (General Education Code(s): W satisfied by taking this course and course 183L.) Prerequisite(s): satisfaction of the Entry Level Writing and Composition requirements; and course 107, 108, or 109; and an undergraduate research contract on file with the department. Concurrent enrollment in course 183L required. D. Potts, B. Sinervo, M. Carr
188. Introduction to Science Writing. S
A rigorous examination and practice of the skills involved in writing articles about science, health, technology, and the environment for the general public. Covers the essential elements of news writing and explanatory journalism, including developing a story idea, interviewing scientists, fact checking, composition, and editing of multiple drafts about scientific research. (Also offered as Science Communication 160. Students cannot receive credit for both courses.) Prerequisite(s): satisfaction of the Entry Level Writing and C1, C2 requirements. Enrollment restricted to junior and senior biological sciences majors. Enrollment limited to 18. (General Education Code(s): W.) R. Irion
190. Senior Seminar (2 credits). S
Satisfies the senior exit requirement for all biological sciences majors. (Also offered as Biology: Molecular Cell & Dev 190. Students cannot receive credit for both courses.) J. Lee
193. Independent Research in EEB. F,W,S
Supervised undergraduate research on a project with an Ecology and Evolutionary Biology (EEB) faculty member, adjunct, or affiliate mentor. Prerequisites: course 183W and an undergraduate research contract on file with the department. M. Carr
193F. Independent Research in EEB (2 credits). *
Supervised undergraduate research on a project with an Ecology and Evolutionary Biology (EEB) faculty member, adjunct, or affiliate mentor. Prerequisites: course 183W and an undergraduate research contract on file with the department. M. Carr
195. Senior Thesis. F,W,S
An individually supervised course, with emphasis on independent research. Students required to submit a senior thesis. Enrollment restricted to majors in biology, ecology and evolution, marine biology, plant sciences, and the combined major with environmental studies. Students submit petition to sponsoring agency. The Staff
198. Independent Field Study. F,W,S
Provides for individual programs of study (a) by means other than the usual supervision in person, or (b) when the student is doing all or most of the course work off campus. With permission of the department, may be repeated for credit, or two or three courses taken concurrently. Students submit petition to sponsoring agency. May be repeated for credit. The Staff
198F. Independent Field Study (2 credits). F,W,S
Provides for two units of independent field study (a) by means other than the usual supervision in person, or (b) when the student is doing all or most of the course work off campus. Students submit petition to sponsoring agency. May be repeated for credit. The Staff
199. Tutorial. F,W,S
Reading, discussion, written reports, and laboratory research on selected biological topics, using facilities normally available on campus. Students submit petition to sponsoring agency. May be repeated for credit. The Staff
199F. Tutorial (2 credits). F,W,S
Two-unit Tutorial. Reading, discussion, written reports, and laboratory research on selected biological topics, using facilities normally available on campus. Students submit petition to sponsoring agency. May be repeated for credit. The Staff
Graduate Courses
200A. Scientific Skills. F
Exposes graduate students to teaching skills, understanding the scientific method, searching and organizing literature, grant proposal and scientific writing, data management and presentation, and scientific speaking. Students are evaluated on their participation and the quality of a written research proposal. Enrollment restricted to graduate students. M. Carr
200B. Advanced Organismal Biology. F
Consists of lectures focusing on pivotal topics in ecology and evolution. Relevant background material is developed followed by a critical analysis of readings from the primary literature. Designed to give graduate (and advanced undergraduate) students direct contact with the major areas of research that are currently at the forefront of organismal biology. Enrollment restricted to graduate students. B. Lyon, J. Estes, D. Costa
208. Marine Ecology. W
Paradigms and designs in marine ecology. A review of the paradigms that have shaped our understanding of marine ecology; analysis and discussion of experiments with these paradigms. Students cannot receive credit for this course and course 108. Enrollment restricted to graduate students. M. Carr
245. Plant Ecology. F
An exploration of the ecology of plant form, function, distribution, abundance, and diversity. Topics include plant adaptations to environmental conditions, life history variation, competition, reproductive ecology, herbivory, and patterns of diversity. Lecture with discussions of original papers and independent field project. Students cannot receive credit for this course and course 145. Prerequisite(s): BIOE 107 or ENVS 24 or permission of instructor. Concurrent enrollment in BIOE 245L is required except by permission of instructor. Enrollment restricted to graduate students. I. Parker
245L. Field Methods in Plant Ecology Laboratory. F
Hands-on exploration of the concepts and techniques of plant ecology. A combination of lab, greenhouse, and field-based exercises (irrespective of weather conditions), statistical analysis, and scientific writing. One required weekend field trip. Students cannot receive credit for this course and course 145L. Concurrent enrollment in BIOE 245 is required. Enrollment restricted to graduate students. Enrollment limited to 2. I. Parker
247. Community Ecology. S
Develops the major themes of community ecology: structure, trophic dynamics, succession, complex interactions among species, herbivory, evolution, and coevolution. Uses case histories of well-studied marine and terrestrial systems. Students cannot receive credit for this course and course 147. Enrollment restricted to graduate students. L. Fox
258L. Experimental Marine Ecology. *
Supervised individual research projects in experimental marine biology. Students carry out a complete research project, including (1) the formation of hypotheses, (2) the design and implementation of experiments, (3) collection, analysis, and interpretation of data, and (4) the write-up of an oral presentation. Prerequisite(s): BIOE 208; and interview to assess ability to carry out field project. Enrollment limited to 20. Offered in alternate academic years. M. Carr, P. Raimondi
272. Population Genetics. F
Basic population genetics and selected topics are covered including genetics of speciation, tempo and mode of evolution, genetics of social behavior, natural selection in human populations, and the impact of molecular studies on evolutionary theory. Students cannot receive credit for this course and Biology 172. Concurrent enrollment in BIOE 272L is required. Enrollment restricted to graduate students. Offered in alternate academic years. G. Pogson
272L. Population Genetics Laboratory (2 credits). F
A companion course to 272, Population Genetics, that applies the theory developed in that course to related disciplines including conservation biology, ecology, agriculture, and population biology. Original scientific literature relating to the theory developed in course 272 is read, and applied problem sets are solved by the students. Students cannot receive credit for this course and course 172L. Must be taken concurrently with BIOE 272. Enrollment restricted to graduate students. Offered in alternate academic years. G. Pogson
274. Evolutionary Game Theory. W
Reviews static equilibrium concepts, games of incomplete information, and the traditional theory of dynamic games in discrete time. Develops recent evolutionary game models, including replicator and best reply dynamics, and applications to economics, computer science, and biology. Prerequisite(s): upper-division math courses in probability theory are strongly recommended. Cannot receive credit for this course and Economics 166B or Computer Science 166B. (Also offered as Computer Science 272. Students cannot receive credit for both courses.) B. Sinervo, The Staff
279. Evolutionary Ecology. W
Analysis of the ways in which ongoing evolution and coevolution shape the ecological structure and dynamics of populations, species, and species interactions across geographic landscapes. Enrollment restricted to graduate students. The Staff
281A. Topics in Basic and Applied Marine Ecology. F,W,S
Seminar focusing on concepts in basic and applied ecology. Structure rotates quarterly between graduate student research and readings of journal articles and textbooks. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission of instructor. Enrollment limited to 10. M. Carr
281B. Topics in Molecular Evolution (2 credits). F,W,S
A discussion of current research and literature review on the subject of molecular evolution. Primary focus on recent results on molecular phylogenetics and molecular population genetics. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission of instructor. May be repeated for credit. G. Bernardi
281C. Topics in Physiological Ecology. F,W,S
An intensive seminar focusing on the interaction between physiological constraint and life history options and solutions employed by animals. Topics vary from comparative physiology to ecological theory. Participants are required to present results of their own research or review papers of interest. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission of instructor. May be repeated for credit. D. Costa
281E. Topics in Freshwater Ecology. F,W,S
Current topics in freshwater ecology, eco-evolutionary dynamics, fisheries, and fish ecology. Enrollment restricted to graduate students. Qualified undergraduates may enroll with permission from instructor. Enrollment limited to 18. May be repeated for credit. E. Palkovacs
281F. Ecological Research Topics. F,W,S
Intensive research and discussions on plant-animal interactions. All students undertake a research project and meet weekly with the faculty sponsor to monitor progress. The group meets weekly to discuss experimental design and analysis, specific problems related to the students' research, relevant research papers, or manuscripts that the group members are writing. Each student gives a formal presentation of research plans or progress each quarter. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission of instructor. Enrollment limited to 10. May be repeated for credit. L. Fox
281H. Topics in Comparative Marine Physiology. F,W,S
Intensive seminar on selected topics in marine physiology. Students present results from their own research and discuss recent advances from the literature. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission of instructor. May be repeated for credit. R. Mehta
281I. Topics in Disease Ecology, Population Biology, and Conservation. F,W,S
Selected topics in population biology and disease ecology. Students present results from their own research and discuss recent advances from the literature. (Formerly "Topics in Plant Population and Disease Ecology") Enrollment restricted to graduate students; qualified undergraduates may enroll by permission of instructor. Enrollment limited to 18. May be repeated for credit. A. Kilpatrick
281J. Topics in Research on Biochemical Ecology. *
Seminar in which students give critically evaluated presentations regarding current research on selected topics in plant ecology with an emphasis on biochemical ecology. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission of instructor. Enrollment limited to 12. May be repeated for credit. J. Langenheim
281K. Topics in Plant Evolution. F,W,S
Intensive seminar on selected topics in plant evolution. Students present results from their own research and discuss recent advances from the literature. Enrollment restricted to graduate students; qualified undergraduates may enroll by permission of instructor. Enrollment limited to 18. May be repeated for credit. K. Kay
281L. Topics in Behavioral and Evolutionary Ecology. F,W,S
An intensive seminar on selected topics in behavioral and evolutionary ecology. Students are expected to discuss the current literature and present literature reviews, research proposals, and preliminary results from their ongoing research. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission of instructor. Enrollment limited to 10. B. Lyon
281N. Topics in Marine Vertebrate Ecology. F,W,S
Seminar on the ecology of marine vertebrates. Topics vary from the factors that explain the distribution of marine predators to island biogeography and the ecosystem effects of introduced vertebrates on islands. Enrollment restricted to graduate students. Enrollment limited to 12. May be repeated for credit. D. Croll
281O. Topics in Plant-Water Relations. F,W,S
Intensive seminar focusing on fundamental and evolutionary concepts in plant-water relations. Students present results from their own research and discuss recent advances from the literature. Enrollment restricted to graduate students; qualified undergraduates may enroll by permission of instructor. Enrollment limited to 18. May be repeated for credit. J. Pittermann
281P. Topics in Plant Population Ecology. F,W,S
An intensive seminar on selected topics in plant ecology and population biology. Students present results from their own research and discuss recent advances from the literature. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission from instructor. Enrollment limited to 12. May be repeated for credit. I. Parker
281Q. Topics in Molecular Evolutionary Genetics. F,W,S
An intensive seminar on selected topics in molecular evolutionary genetics. Students are required to present results from their own research projects, present a critical review paper at least once during the quarter, and submit a written research proposal. Enrollment restricted to graduate students; qualified undergraduate students may enroll with permission of instructor. Enrollment limited to 10. May be repeated for credit. G. Pogson
281R. Topics in Marine Ecology and Evolutionary Biology. F,W,S
An intensive seminar series focusing on fundamental concepts in marine ecology. Emphasis changes quarter to quarter. At least one quarter per year is devoted to discussion of graduate student research. Other quarters involve reading and evaluating current and classic literature on marine ecology and evolutionary biology. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission of instructor. Enrollment limited to 10. May be repeated for credit. P. Raimondi
281S. Topics in Ancient DNA and Paleogenomics. F,W,S
Topics in population genetics and genomics, focusing on work involving paleontological and archaeological material. Students present weekly written and oral reports of their research projects. Once each term, students critique a recent publication. Enrollment restricted to graduate students. Qualified undergraduates may enroll with permission from instructor. May be repeated for credit. B. Shapiro
281T. Species Interactions and Coevolution. F,W,S
The genetics and ecological structure of species interactions, and the role of coevolution between species in shaping biodiversity. Enrollment restricted to graduate students. Enrollment limited to 15. May be repeated for credit. J. Thompson
281U. Topics in Invertebrate Biology. F,W,S
An intensive study about concepts, theory, and techniques for graduate students conducting research on the ecology, genetics, evolution, systematics, or biodiversity of marine invertebrates. Enrollment restricted to graduate students; advanced undergraduates may enroll with permission of instructor. Enrollment limited to 15. May be repeated for credit. D. Potts
281V. Topics in Behavioral Ecology. F,W,S
A discussion of current topics and methods in behavioral ecology and life history evolution. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission of instructor. May be repeated for credit. B. Sinervo
281W. Topics in Exercise and Environmental Physiology. F,W,S
A weekly seminar discussion on current research and techniques in mammalian exercise and environmental physiology. Areas covered include locomotor physiology, exercise testing and cardiovascular monitoring, and biomechanics. Oral presentation of ongoing research or current literature required from each student. Enrollment restricted to graduate students; qualified undergraduates may enroll with permission of instructor. May be repeated for credit. T. Williams
286. Experimental Design and Data Analysis. W
Focuses on problems and designs in ecology and population biology. Topics include basic experimental design; exploratory data analysis—from a graphical perspective; hands-on statistics; and graphical theory. Structured around a statistical analysis and graphics computer program to teach students to design their own surveys and experiments and analyze their data correctly. Students cannot receive credit for this course and course 186. Prerequisite(s): one course in statistics or by permission of instructor. Enrollment restricted to graduate students. P. Raimondi
286L. Experimental Design and Data Analysis Lab (2 credits). W
Required lab that accompanies Biology 286. Lab will focus on hands-on statistical problem solving, graphical presentations and experimental design issues. Concurrent enrollment in course 286 is required. Enrollment restricted to graduate students. P. Raimondi
293. Readings in Ecology and Evolution (2 credits). W,S
Weekly readings and discussions of recent research papers in ecology, evolution, and related topics from organismal biology. Enrollment restricted to graduate students. May be repeated for credit. A. Kilpatrick, L. Fox
294. Ecology, Evolutionary Biology Seminar (no credit). F,W,S
Selected topics of current interest to ecologists and evolutionary biologists presented by weekly guest speakers. Enrollment restricted to graduate students. T. Williams, B. Shapiro, D. Croll
295. Advanced Ecology and Evolutionary Biology Seminar (no credit). F,W,S
Course consists of extended weekly meetings organized around an advanced theme in theoretical or applied evolutionary biology, ecology, physiology, behavior, or other aspect of oranismal biology. Course is targeted at students who already have reached a professional level of expertise in their field and advanced master students. Enrollment restricted to graduate students. Enrollment limited to 24. The Staff
297. Independent Study. F,W,S
Independent study for graduate students who have not yet settled on a research area for their thesis. Students submit petition to sponsoring agency. The Staff
299. Thesis Research. F,W,S
Students submit petition to sponsoring agency. The Staff
Revised: 09/01/14
